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Dorastający młodzian w ramach eksperymentu zaczyna uczęszczać do szkoły pełnej dorastających niewiast, którym do tej pory nie było dane obcować z płcią przeciwną.

The traditional girls-only Yuihime Private Academy is considering the possibility of starting to accept both genders and has thus made an agreement with a neighboring school to receive some of its students as an experiment to see the reaction to this adaptation. Uryuu Shingo is one of the many students transferred for a ten month experimental phase at Yuihime Academy. He, his sister, and his fellow relocated colleagues are impressed by the sophisticated campus, but even more by the resistance of many students who don't want the boys over there.Source: ANN

The traditional girls-only Yuihime Private Academy is considering the possibility of starting to accept both genders and has thus made an agreement with a neighboring school to receive some of its students as an experiment to see the reaction to this adaptation. Shingo Iryu is one of the many students transferred for a ten month experimental phase at Yuihime Academy. He, his sister, and his fellow relocated colleagues are impressed by the sophisticated campus, but even more by the resistance of many students who don't want the boys over there.

One day, Shingo's school combines with a girls' school. As a specially selected student, Shingo temporarily joins the girls' school until they officially consolidate. He meets various unique girls there, but they all dislike male students. To improve their relationship, Shingo works hard, and...

The story of Mashiroiro Symphony primarily takes place in the fictional town of Kagamidai (各務台?), which consists of two distinct districts dubbed the "Old District" (旧市街, Kyū-shigai?), which primarily contains traditional housings resided by upper class families, and the "New District" (新市街, Shin-shigai?), which is generally inhabited by working class families. The town also houses two private academic institutions undergoing a merger: the Yuihime Girls' Private Academy (私立結姫女子学園, Shiritsu Yuihime Joshi Gakuen?), a notorious upper class girls' academy nicknamed Yuijo (結女?), and the Kagamidai Private Academy (私立各務台学園, Shiritsu Kagamidai Gakuen?), a coeducation school which Shingo, the protagonist whose role the player assumes, attends. As part of the merger, students from both schools are selected to participate in a test class at the Yuihime Girls' Academy campus,[1] an act which is initially opposed by the female student body.